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The R groups that can be protonated/deprotonated include Lys, Arg, and His, which have a + 1 charge when protonated, and Glu and Asp (carboxylic acids), Tyr and Ser (alcohols), and Cys (thiol), which have zero charges when protonated

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Exp Cell Res 306:18 Piotrowicz K, Gasowski J, Michel JP, Veronese N (2021) Post-Covid-19 acute sarcopenia: physiopathology and management

[DOI] [PMC free article] [PubMed] [Google Scholar] Celli A, Que FG, Gores GJ, LaRusso NF

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